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Tinbergen's four questions : ウィキペディア英語版
Tinbergen's four questions
Tinbergen's four questions, named after Nikolaas Tinbergen, are complementary categories of explanations for behaviour. It suggests that an integrative understanding of behaviour must include both a proximate and ultimate (functional) analysis of behaviour, as well as an understanding of both phylogenetic/developmental history and the operation of current mechanisms.〔Daly, M & Wilson, M. (1983). Sex, evolution, and behaviour. Brooks-Cole.〕
==Four categories of questions and explanations==

When asked about the purpose of sight in humans and animals, even elementary school children can answer that animals have vision to help them find food and avoid danger (adaptation). Biologists have three additional explanations: sight is caused by a particular series of evolutionary steps (phylogeny), the mechanics of the eye (causation), and even the process of an individual’s development (ontogeny). Although these answers may be very different, they are consistent with each other. This idea was formulated in the 1960s when Tinbergen delineated the four questions based on Aristotle's Four Causes.〔Hladký, V. & Havlíček, J. (2013). ''(Was Tinbergen an Aristotelian? Comparison of Tinbergen’s Four Whys and Aristotle’s Four Causes )''. ''Human Ethology Bulletin'', 28(4), 3-11〕 This schema constitutes a basic framework of the overlapping behavioural fields of ethology & anthropology, behavioural ecology, sociobiology & evolutionary psychology, and comparative psychology. It was in fact Julian Huxley who identified the first three questions, Niko Tinbergen gave only the fourth question, but Julian Huxley's questions failed to distinguish between survival value and evolutionary history, so Tinbergen's fourth question helped resolve this problem.〔p411 in Tinbergen, Niko (1963) "On Aims and Methods in Ethology," Zeitschrift für Tierpsychologie, 20: 410–433.〕

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